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Most krill species display large daily vertical migrations , thus providing food for predators near the surface at night and in deeper waters during the day.
Krill are fished commercially in the Southern Ocean and in the waters around Japan. Most of the krill catch is used for aquaculture and aquarium feeds, as bait in sport fishing, or in the pharmaceutical industry.
They are eaten as camarones in Spain and Philippines. In the Philippines, krill are also known as alamang and are used to make a salty paste called bagoong.
Krill are also the main prey of baleen whales , including the blue whale. Krill belong to the large arthropod subphylum , the Crustacea.
The most familiar and largest group of crustaceans, the class Malacostraca , includes the superorder Eucarida comprising the three orders, Euphausiacea krill , Decapoda shrimp, prawns, lobsters, crabs , and the planktonic Amphionidacea.
The order Euphausiacea comprises two families. The more abundant Euphausiidae contains 10 different genera with a total of 85 species.
Of these, the genus Euphausia is the largest, with 31 species. It is considered the most primitive extant krill species. Well-known species of the Euphausiidae of commercial krill fisheries include Antarctic krill Euphausia superba , Pacific krill E.
As of [update] , the order Euphausiacea is believed to be monophyletic due to several unique conserved morphological characteristics autapomorphy such as its naked filamentous gills and thin thoracopods  and by molecular studies.
There have been many theories of the location of the order Euphausiacea. Since the first description of Thysanopode tricuspide by Henri Milne-Edwards in , the similarity of their biramous thoracopods had led zoologists to group euphausiids and Mysidacea in the order Schizopoda , which was split by Johan Erik Vesti Boas in into two separate orders.
Molecular studies have not unambiguously grouped them, possibly due to the paucity of key rare species such as Bentheuphausia amblyops in krill and Amphionides reynaudii in Eucarida.
One study supports the monophyly of Eucarida with basal Mysida ,  another groups Euphausiacea with Mysida the Schizopoda ,  while yet another groups Euphausiacea with Hoplocarida.
No extant fossil can be unequivocally assigned to Euphausiacea. Some extinct eumalacostracan taxa have been thought to be euphausiaceans such as Anthracophausia , Crangopsis — now assigned to the Aeschronectida Hoplocarida  — and Palaeomysis.
Krill occur worldwide in all oceans, although many individual species have endemic or neritic i. Bentheuphausia amblyops , a bathypelagic species, has a cosmopolitan distribution within its deep-sea habitat.
Species of the genus Thysanoessa occur in both Atlantic and Pacific oceans. Northern krill occur across the Atlantic from the Mediterranean Sea northward.
Species with neritic distributions include the four species of the genus Nyctiphanes. Species with endemic distributions include Nyctiphanes capensis , which occurs only in the Benguela current,  E.
In the Antarctic, seven species are known,  one in genus Thysanoessa T. Other species known in the Southern Ocean are E.
Krill are crustaceans and have a chitinous exoskeleton made up of three tagmata: This outer shell of krill is transparent in most species.
Krill feature intricate compound eyes ; some species adapt to different lighting conditions through the use of screening pigments. These thoracic legs include feeding legs and grooming legs.
Additionally all species have five pairs of swimming legs called pleopods or "swimmerets", very similar to those of a lobster or freshwater crayfish.
Most krill are about 1—2 centimetres 0. The largest krill species is the bathypelagic Thysanopoda spinicauda. Except for Bentheuphausia amblyops , krill are bioluminescent animals having organs called photophores that can emit light.
The light is generated by an enzyme -catalysed chemiluminescence reaction, wherein a luciferin a kind of pigment is activated by a luciferase enzyme.
Studies indicate that the luciferin of many krill species is a fluorescent tetrapyrrole similar but not identical to dinoflagellate luciferin  and that the krill probably do not produce this substance themselves but acquire it as part of their diet, which contains dinoflagellates.
Many krill are filter feeders: These filters can be very fine indeed in those species such as Euphausia spp. Krill are mostly omnivorous ,  although a few species are carnivorous , preying on small zooplankton and fish larvae.
Krill are an important element of the aquatic food chain. Krill convert the primary production of their prey into a form suitable for consumption by larger animals that cannot feed directly on the minuscule algae.
Northern krill and some other species have a relatively small filtering basket and actively hunt copepods and larger zooplankton. Many animals feed on krill, ranging from smaller animals like fish or penguins to larger ones like seals and baleen whales.
Disturbances of an ecosystem resulting in a decline in the krill population can have far-reaching effects. During a coccolithophore bloom in the Bering Sea in ,  for instance, the diatom concentration dropped in the affected area.
Krill cannot feed on the smaller coccolithophores, and consequently the krill population mainly E. This in turn affected other species: The incident was thought to have been one reason salmon did not spawn that season.
Climate change poses another threat to krill populations. Such diseases were reported for Thysanoessa inermis in the Bering Sea and also for E.
The life cycle of krill is relatively well understood, despite minor variations in detail from species to species. The pseudometanauplius stage is exclusive to species that lay their eggs within an ovigerous sac: The larvae grow and moult repeatedly as they develop, replacing their rigid exoskeleton when it becomes too small.
Smaller animals moult more frequently than larger ones. Yolk reserves within their body nourish the larvae through metanauplius stage.
By the calyptopsis stages differentiation has progressed far enough for them to develop a mouth and a digestive tract, and they begin to eat phytoplankton.
By that time their yolk reserves are exhausted and the larvae must have reached the photic zone , the upper layers of the ocean where algae flourish.
During the furcilia stages, segments with pairs of swimmerets are added, beginning at the frontmost segments. Each new pair becomes functional only at the next moult.
The number of segments added during any one of the furcilia stages may vary even within one species depending on environmental conditions.
Krill employ two types of spawning mechanism. These species generally hatch in the nauplius 1 stage, but have recently been discovered to hatch sometimes as metanauplius or even as calyptopis stages.
Moulting occurs whenever a specimen outgrows its rigid exoskeleton. Young animals, growing faster, moult more often than older and larger ones.
The frequency of moulting varies widely by species and is, even within one species, subject to many external factors such as latitude, water temperature, and food availability.
The subtropical species Nyctiphanes simplex , for instance, has an overall inter-moult period of two to seven days: Shrinkage has been postulated for other temperate-zone species of krill as well.
Some high-latitude species of krill can live for more than six years e. Most krill are swarming animals; the sizes and densities of such swarms vary by species and region.
For Euphausia superba , swarms reach 10, to 60, individuals per cubic meter. In , Gandomi and Alavi presented what appears to be a successful stochastic algorithm for modelling the behaviour of krill swarms.
The algorithm is based on three main factors: Krill typically follow a diurnal vertical migration. It has been assumed that they spend the day at greater depths and rise during the night toward the surface.
The deeper they go, the more they reduce their activity,  apparently to reduce encounters with predators and to conserve energy.
Swimming activity in krill varies with stomach fullness. Sated animals that had been feeding at the surface swim less actively and therefore sink below the mixed layer.
Krill with empty stomachs swim more actively and thus head towards the surface. Vertical migration may be a 2—3 times daily occurrence.
Experimental studies using Artemia salina as a model suggest that the vertical migrations of krill several hundreds of metres, in groups tens of metres deep, could collectively create enough downward jets of water to have a significant effect on ocean mixing.
Dense swarms can elicit a feeding frenzy among fish, birds and mammal predators, especially near the surface. When disturbed, a swarm scatters, and some individuals have even been observed to moult instantaneously, leaving the exuvia behind as a decoy.
Their larger migrations are subject to ocean currents. When in danger, they show an escape reaction called lobstering — flicking their caudal structures, the telson and the uropods , they move backwards through the water relatively quickly, achieving speeds in the range of 10 to 27 body lengths per second, which for large krill such as E.
Larval forms of krill are generally considered zooplankton. Krill have been harvested as a food source for humans and domesticated animals since at least the 19th century, and possibly earlier in Japan, where it was known as okiami.
Large-scale fishing developed in the late s and early s, and now occurs only in Antarctic waters and in the seas around Japan. In , two events caused a decline in krill fishing: After an October review, the Commission decided not to change the quota.
Although krill are found worldwide, fishing in Southern Oceans are preferred because the krill are more "catchable" and abundant in these regions.
Particularly in Antarctic seas which are considered as pristine , they are considered a "clean product". In it was announced that almost every krill fishing company operating in Antarctica will abandon operations abandoned in huge areas around the Antarctic Peninsula from , including "buffer zones" around breeding colonies of penguins.
Krill is a rich source of protein and omega-3 fatty acids which are under development in the early 21st Century as human food, dietary supplements as oil capsules, livestock food, and pet food.
For mass-consumption and commercially prepared products, they must be peeled to remove the inedible exoskeleton.
In , the US Food and Drug Administration published a letter of no objection for a manufactured krill oil product to be generally recognized as safe GRAS for human consumption.
The hunter looks up and takes a defensive position, seeing the massive black wyvern perched on a ledge with its evil yellow eyes. The Skrill snarls at the hunter, then spreads its wings and lets out a screech, and lightning begins to course over its body and shoot in random directions.
The Skrill finishes the display and sees the hunter getting up off the ground. It rises into the air and dives, electricity still crackling across its body as it skims the ground and narrowly misses the hunter again.
It lands and lets out its roar as the hunter faces it, ready to begin the hunt. In the foggy Area 5, a Night Fury curls up on the ground to sleep.
The view shifts to the massive skeleton shadowing the area, where a pair of yellow eyes glares down at the black Elder Dragon. A dark shape scuttles up a rib bone and prepares to spring.
Meanwhile, the hunter has just entered Area 5 and immediately catches sight of the snoring Night Fury. Revealing itself to be a massive Skrill, the beast allows a current of electricity to run across its crest, and it flares out its wings intimidatingly.
The Night Fury tries to run, but the Skrill backs it into a corner. The Skrill is a dragon-like beast that has been classified as a Pseudowyvern.
Unfortunately, its wide range places a large number of people and monsters at risk, because the Skrill is so ridiculously aggressive that it is actually considered evil.
Something to note is that the Skrill has an intense hatred of its closest cousin, the Night Fury. Both monsters are very similar, which would explain their animosity toward one another.
But while Night Furies try to leave their enemy alone, a Skrill will ignore all other threats when a Night Fury is detected and try to slay it.
Alongside several other dragon-like monsters, the Skrill initially lived up north before migrating into the various subregions of the known world.
The Skrill can be carved three times, have its tail severed, have its head, back, and both wings broken, and drop an item.
Hunters nearby will have their health drained if they approach too closely. Starts to run forward, then leaps at the hunter in a series of large pounces, similarly to Barioth.
Stands up and flares its wings, then suddenly rockets forward with wings outstretched. It will then turn around and either remain in the air or land.
While in the air, rears up and flaps its wings once, sending a small wave of electricity flying forward a short distance. While in the air, draws a wing back and then swings it in a forward arc while it sparks with electricity.
Throws its head way back and swings it to the left, then lowers it and shoots a bolt of lightning while sweeping it to the right.